The analysis yielded two most-parsimonious trees, 239 steps long with a consistency index (CI) of 0.4017, a retention index (RI) of 0.7534, and a rescaled consistency index (RC) of 0.3026. A strict consensus tree of the two most-parsimonious trees is given in Figure 1. The only difference in topology of the two trees is the relative position of Progeryon . In tree one, Progeryon is the earliest-derived taxon within the 'Goneplacidae' + Hexapodidae clade, and in tree two, it is the sister to the Goneplacidae + Hexa-podidae + Portunoidea clade. The relationships among the major groups, indicating Bremer support and unambiguous character state changes, are given in Figure 2.